Read, G.; Fauchald, K. (Ed.) (2026). World Polychaeta Database. Harmothoe imbricata (Linnaeus, 1767). Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=130769 on 2026-03-21
accepted
Valid name (ICZN) or name considered to be taxonomically correct (ICBN)
Species
Les annélides polychètes des côtes de Dinard. Seconde partie.
Saint-Joseph, Arthur d'Anthoine de. (1888). Les annélides polychètes des côtes de Dinard. Seconde partie. Annales des Sciences Naturelles, Zoologie et Paléontologie, Paris. Série 7, 5(2): 141-338, plates VI-XIII.
Two new species of polychaete worms of the family Polynoidae from Puget Sound and San Juan Archipelago
Pettibone, Marian H. 1948. Two new species of polychaete worms of the family Polynoidae from Puget Sound and San Juan Archipelago. Journal of the Washington Academy of Sciences, 38(12): 412-414.
Contributions towards a monograph of the species of Annelides belonging to the Aphroditacea, containing a list of the known species, and a description of some new species contained in the National Collection of the British Museum.
Baird, William. (1865). Contributions towards a monograph of the species of Annelides belonging to the Aphroditacea, containing a list of the known species, and a description of some new species contained in the National Collection of the British Museum. The Journal of the Linnean Society of London. Zoology. 8(31-32): 172-202.
Polychaetous annelids collected by Captain Robert
Treadwell, Aaron L. 1937. Polychaetous annelids collected by Captain Robert A. Bartlett in Greenland, Fox Basin, and Labrador. Journal of the Washington Academy of Sciences, 27(1): 23-36.
Örsted, Anders Sandoe. (1843). Grönlands Annulata dorsibranchiata. Det Kongelige Danske videnskabernes selskabs. Naturvidenskabelige og mathematiske afhandlinger. 10 (series 4): 153-216. 8 plates.
Histoire naturelle des Annelés marins et d'eau douce. Annélides et Géphyriens
Quatrefages, A. de. (1866 (1865)). Histoire naturelle des Annelés marins et d'eau douce. Annélides et Géphyriens. Librarie Encyclopédique de Roret. Paris. Volume 1. 1-588.
Dopolnenia k faune Annelid Chernogo morya [Additions/supplement to the annelid fauna of the Black Sea]
Bobretzky, N. (1881). Dopolnenia k faune Annelid Chernogo morya [Additions/supplement to the annelid fauna of the Black Sea]. Zapiski Kievskago obshchestva estestvoispytateleĭ [Mémoires de la Société des Naturalistes de Kiew]. 6(2): 183-212, plates 6 & 7.
Miscellanea zoologica. Quibus novae imprimis atque obscurae animalium species describuntur et observationibus iconibusque illustrantur. Petrum van Cleef
Pallas, P.S. (1766) Miscellanea zoologica. Quibus novae imprimis atque obscurae animalium species describuntur et observationibus iconibusque illustrantur. Petrum van Cleef. Hagí Comitum., xii + 224 pp.;14 pls.
Systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Ed. 12. 1., Regnum Animale. 1 & 2. [The system of nature through the three kingdoms of nature: according to classes, orders, genera, species, with characters, differences, synonyms, places. Ed. 12. 1., Animal Kingdom. 1 & 2]
Linnaeus, C. (1767). Systema naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Ed. 12. 1., Regnum Animale. 1 & 2. [The system of nature through the three kingdoms of nature: according to classes, orders, genera, species, with characters, differences, synonyms, places. Ed. 12. 1., Animal Kingdom. 1 & 2]. Holmiae [Stockholm], Laurentii Salvii. pp. 1-532 [1766] pp. 533-1327 [1767].
The Powers of the Creator Displayed in the Creation: or, Observations on Life Amidst the Various Forms of the Humbler Tribes of Animated Nature: with Practical Comments and Illustrations. Volume II.
Dalyell, J. G. (1853). The Powers of the Creator Displayed in the Creation: or, Observations on Life Amidst the Various Forms of the Humbler Tribes of Animated Nature: with Practical Comments and Illustrations. Volume II. xiii + 359 pp., 46 pls. John van Voorst. London.
Zoologiae Danicae Prodromus, seu Animalium Daniae et Norvegiae Indigenarum characters, nomina, et synonyma imprimis popularium. [Prodromus of Danish Zoology, or the characters, names, and synonyms of the indigenous animals of Denmark and Norway, especially of the popular ones.]
Müller, O.F. (1776). Zoologiae Danicae Prodromus, seu Animalium Daniae et Norvegiae Indigenarum characters, nomina, et synonyma imprimis popularium. [Prodromus of Danish Zoology, or the characters, names, and synonyms of the indigenous animals of Denmark and Norway, especially of the popular ones.]. Typis Hallageriis, Havniae. 32: 1-282.
Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus differentiis, synonymis, locis. [The system of nature through the three kingdoms of nature, according to classes, orders, genera, species, with characteristic differences, synonyms, and places.]
Linnaeus, C. (1788). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus differentiis, synonymis, locis. [The system of nature through the three kingdoms of nature, according to classes, orders, genera, species, with characteristic differences, synonyms, and places.]. Editio decima tertia, aucta reformata. Cura J.F. Gmelin. G.E. Beer, Lipsiae. 1(5-6):2225-3909.
Beskrivelse over Norske Insecter. Andet Stykke.
Strøm H. (1768). Beskrivelse over Norske Insecter. Andet Stykke. Det Kongelige Norske Videnskabers Selskabs Skrifter, Kjobenhavn. 4: 313-371, pl. 16.
British Zoology, vol. IV. Crustacea. Mollusca. Testacea.
Pennant, T. (1777). British Zoology, vol. IV. Crustacea. Mollusca. Testacea. London. i-viii, 1-154, pls. 1-93.
Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Polychaeta
Budaeva N.E., Jirkov I.A., Savilova T.A., Paterson G.L.J. (2014). Deep-sea fauna of European seas: An annotated species check-list of benthic invertebrates living deeper than 2000 m in the seas bordering Europe. Polychaeta. Invertebrate Zoology. Vol.11. No.1: 217–230 [in English].
BU. (2013). Provision of services for field sampling, species identification and data analysis of benthic faunal communities of Hong Kong marin waters. Final report submitted to EPD.
Monitoring van de effecten van de verruiming 48'/43': MOVE eindrapport 2006 (Rapport 10)
van Eck, G. Th. M.; Holzhauer, H. (2007). Monitoring van de effecten van de verruiming 48'/43': MOVE eindrapport 2006 (Rapport 10). Rapport RIKZ = Report RIKZ, 2007.003. RIKZ: Middelburg. 64 pp.
Annulata Polychaeta Spetsbergiæ, Grœnlandiæ, Islandiæ et Scandinaviæ. Hactenus Cognita.
Malmgren, A.J. (1867). Annulata Polychaeta Spetsbergiæ, Grœnlandiæ, Islandiæ et Scandinaviæ. Hactenus Cognita. Ex Officina Frenckelliana, Helsingforslæ. 127 pp. & XIV plates.
A monograph of British Annelids. vol.1. pt.2. Polychaeta Amphinomidae to Sigalionidae
McIntosh, W.C. 1900. A monograph of British Annelids. vol.1. pt.2. Polychaeta Amphinomidae to Sigalionidae. Ray Society of London, 1(l2): 215-442.
Marine polychaete worms of the New England region. I. Aphroditidae through Trochochaetidae
Pettibone, Marian H. (1963). Marine polychaete worms of the New England region. I. Aphroditidae through Trochochaetidae. Bulletin of the United States National Museum. 227(1): 1-356.
Colour polymorphism in the polychaete Harmothoe imbricata (Linnaeus, 1767)
Nygren, Arne; Norlinder, Erika; Panova, Marina ; Pleijel, Fredrik. (2011). Colour polymorphism in the polychaete Harmothoe imbricata (Linnaeus, 1767). Marine Biology Research. 7(1): 54 - 62.
[Checklist of marine biota of China seas]
Liu, J.Y. [Ruiyu] (ed.). (2008). [Checklist of marine biota of China seas]. China Science Press. 1267 pp.
Cobscook Bay inventory: a historical checklist of marine invertebrates spanning 162 years
Trott, T. J. (2004). Cobscook Bay inventory: a historical checklist of marine invertebrates spanning 162 years. Northeastern Naturalist. 11, 261-324.
Tri-oceanic connectivity of the supposedly cosmopolitan polychaete, Harmothoe imbricata (Annelida: Polynoidae): insights from the COI marker
David, Andrew A.; Cahill, Jordan. (2020). Tri-oceanic connectivity of the supposedly cosmopolitan polychaete, Harmothoe imbricata (Annelida: Polynoidae): insights from the COI marker. Marine Biology Research. 16(4): 256-264.
Annelida, Borstenwürmer, Polychaeta [Annelida, bristleworms, Polychaeta]
Hartmann-Schröder, G. (1996). Annelida, Borstenwürmer, Polychaeta [Annelida, bristleworms, Polychaeta]. 2nd revised ed. The fauna of Germany and adjacent seas with their characteristics and ecology, 58. Gustav Fischer: Jena, Germany. ISBN 3-437-35038-2. 648 pp.
Catalogue of the marine invertebrates of the estuary and Gulf of St. Lawrence.
Brunel, P., L. Bosse & G. Lamarche. (1998). Catalogue of the marine invertebrates of the estuary and Gulf of St. Lawrence. Canadian Special Publication of Fisheries and Aquatic Sciences, 126. 405 pp.
A Practical Guide to the Marine Animals of Northeastern North America.
Pollock, L.W. (1998). A practical guide to the marine animals of northeastern North America. Rutgers University Press. New Brunswick, New Jersey & London. 367 pp.
The Second South Red Sea Expedition, 1965, Report No. 9. Quelques Annélides Polychètes de la Mer Rouge
Fishelson, L.; Rullier, F. (1969). The Second South Red Sea Expedition, 1965, Report No. 9. Quelques Annélides Polychètes de la Mer Rouge. Israel Journal of Zoology. 18: 49-117.
The types of the polychaete worms of the families Polynoidae and Polyodontidae in the United States National Museum and the description of a new genus
Hartman, Olga. (1938). The types of the polychaete worms of the families Polynoidae and Polyodontidae in the United States National Museum and the description of a new genus. Proceedings of the United States National Museum. 86(3046): 107-134.
Catalogue of the Polychaetous Annelids of the World. Parts 1 and 2
Hartman, Olga. (1959). Catalogue of the Polychaetous Annelids of the World. Parts 1 and 2. Allan Hancock Foundation Occasional Paper. 23: 1-628.
Izuka, Akira. (1912). The errantiate Polychaeta of Japan. Journal of the College of Science, Imperial University of Tokyo. 30(2): 1-262, plates I-XXIV.
A preliminary account of the marine annelids of the Pacific coast, with descriptions of new species
Johnson, Herbert P. 1897. A preliminary account of the marine annelids of the Pacific coast, with descriptions of new species. Proceedings of the California Academy of Sciences, Ser. 3, 1(5): 153-199.
The Polychaeta of the Puget Sound region.
Johnson, Herbert Parlin. (1901). The Polychaeta of the Puget Sound region. Proceedings of the Boston Society for Natural History. 29(18): 381-437, plates 1-19.
A catalogue of the British non-parasitical worms in the collection of the British Museum
Johnston, G. (1865). A catalogue of the British non-parasitical worms in the collection of the British Museum. [book]. 1-365. British Museum. London. [See also separate entry for Baird supplement].
Polychaetous annelids from Sagami Bay and Sagami Sea collected by the Emperor Showa of Japan and deposited at the Showa Memorial Institute, National Science Museum, Tokyo. Families Polynoidae and Acoetidae
Imajima, Minoru. (1997). Polychaetous annelids from Sagami Bay and Sagami Sea collected by the Emperor Showa of Japan and deposited at the Showa Memorial Institute, National Science Museum, Tokyo. Families Polynoidae and Acoetidae. National Science Museum Monographs. 13: 1-131.
[Polychaeta of the Arctic Ocean] (In Russian) Polikhety severnogo Ledovitogo Okeana
Jirkov, I.A. (2001). [Polychaeta of the Arctic Ocean] (In Russian) Polikhety severnogo Ledovitogo Okeana. Yanus-K Press, Moscow, 632 pp.
Ruff, R. Eugene. (1995). Family Polynoidae Malmgren, 1867. 105-166. IN: Blake, James A., Hilbig, Brigitte, and Scott, Paul H. (Ed.). Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel. Volume 5 - The Annelida Part 2. Polychaeta: Phyllodocida (Syllidae and scale-bearing families), Amphinomida, and Eunicida. Santa Barbara Museum of Natural History. Santa Barbara.
Nordiska Hafs-Annulater. [part one of three]
Malmgren, Anders Johan. (1865). Nordiska Hafs-Annulater. [part one of three]. Öfversigt af Königlich Vetenskapsakademiens förhandlingar, Stockholm. 22(1): 51-110, plates VIII-XV.
Polychaetes from Scottish Waters, Part 1. Family Polynoidae.
Tebble, Norman; Chambers, Susan. (1982). Polychaetes from Scottish Waters, Part 1. Family Polynoidae. Royal Scottish Museum Studies. [unnumbered] : 1-73.
Have the eyes of bioluminescent scale worms adapted to see their own light? A comparative study of eyes and vision in Harmothoe imbricata and Lepidonotus squamatus
Garm, Anders; Simonsen, Sidsel H.; Mendoza-González, Paula; Worsaae, Katrine; ;. (2021). Have the eyes of bioluminescent scale worms adapted to see their own light? A comparative study of eyes and vision in Harmothoe imbricata and Lepidonotus squamatus. Journal of Experimental Biology. 224(14): ?-?.
Source: http://www.iopan.gda.pl/projects/biodaff/EMBS-0605.html
Distribution for Harmothoe imbricata incerta (Bobretzky, 1881) - European waters (ERMS scope)
Distribution for Harmothoe imbricata (Linnaeus, 1767) - North West Atlantic
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Baie de la Seine
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Golfe Normanno-Breton
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Dutch Exclusive Economic Zone
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Plymouth
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Roscoff
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Wadden Sea
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Zeeland
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Red Sea
Distribution for Harmothoe imbricata (Linnaeus, 1767) - East Africa
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Ukrainian Exclusive Economic Zone
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Cobscook Bay
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Gulf of Maine
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Belgian Exclusive Economic Zone
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Spuikom
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Greek Exclusive Economic Zone
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Irish Exclusive economic Zone
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Canadian Exclusive Economic Zone [Arctic part]
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Manicouagan
Distribution for Harmothoe imbricata (Linnaeus, 1767) - St. Lawrence Estuary
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Grand Banks
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Saguenay Fjord
Distribution for Harmothoe imbricata (Linnaeus, 1767) - Gulf of St. Lawrence
Distribution for Harmothoe maxillospinosa de Saint-Joseph, 1888 - Roscoff
Distribution for Harmothoe maxillospinosa de Saint-Joseph, 1888 - Golfe Normanno-Breton
Distribution for Harmothoe maxillospinosa de Saint-Joseph, 1888 - Dinard
Distribution for Harmothoe unicolor Baird, 1865 - British Columbia
Distribution for Harmothoe levis Treadwell, 1937 - East Greenland
Distribution for Lepidonote cirrata Örsted, 1843 - Öresund
Distribution for Polynoe incerta Bobretzky, 1881 - Black Sea
Distribution for Aphrodita lepidota Pallas, 1766 - West Europe
Distribution for Aphrodita imbricata Linnaeus, 1767 - West Europe
Distribution for Aphrodita varians Dalyell, 1853 - United Kingdom Exclusive Economic Zone
Distribution for Aphrodita cirrata Müller, 1776 - Danish Exclusive Economic Zone
Distribution for Aphrodita plana Gmelin in Linnaeus, 1788 - West Europe
Distribution for Aphrodita violacea Strøm, 1768 - Norwegian Exclusive Economic Zone
Distribution for Aphrodita lepidota minuta Pennant, 1777 - United Kingdom Exclusive Economic Zone
Taxon Description Body size for Taxon denoted by name 130769
Taxon Description AMBI ecological group for Taxon denoted by name 130769
Taxon Description Functional group for Taxon denoted by name 939
Taxon Description Body size (qualitative) for Taxon denoted by name 939
Taxon Description Nomenclature code for Taxon denoted by name 2
Specification Context: Unit mm
Specification Context: Type minimum
Specification Context: Dimension length
Specification Context: Locality (MRGID) 1920
Specification Context: Type maximum
Specification Context: Life stage adult